OHIO PRAIRIES

A “prairie” is a permanent grassland with few or no woody plants. Prairies are fire-dependent ecosystems. Prairies differ from old fields and meadows in that the prairies are more or less stable and self-perpetuating, rather than being a temporary successional stage on the way to becoming a forest. The vegetation that occurs in prairie is mainly drought-tolerant graminoids (grasses and sedges), the growing parts of which are at ground level or below, and so are not harmed by occasional wildfires. Prairie graminoids are usually accompanied by other herbaceous plants, generally called “forbs,” that are adapted to drought and also low nutrient levels. (“Forb” is the term for any non-woody non-graminoid, i.e., what we’d normally call a “wildflower.”)

Prairies occur world-wide, generally in the interior of continents. North American prairie is drier in the west, bordering desert, and the type of prairie that occurs there is called “shortgrass prairie.” In the moister eastern half, adjoining deciduous forest, there is “tallgrass prairie.” Ohio is not usually thought of as a prairie state. Here are two maps from authoritative publications. Note how one shows us well outside the prairie biome, while the other correctly a few scattered prairie patches well within our boundaries.

Prairie in Ohio

Maps vary,  but there are prairies in Ohio.

Generally, where there is adequate moisture year-round, trees can survive, and so the principal vegetation type in Ohio is forest. It is believed that the prairies in Ohio, which comprised only about 1% of the state (and, sadly, of which only about 1% remain) are remnants of a time between 8,000 and 4000 years ago called the “Xerothermic Interval” when the climate was hotter and drier than today, and prairie extended as an eastward-projecting tounge across present-day Ohio, called the “Prairie Peninsula.” Fires deliberately set as a game-management tool by Native Americans are believed to have helped maintain a few prairies into the present day.Today, fire is still used as a management tool to maintain prairies. The principal benefit of fire is to eliminate woody vegetation, as the above-ground buds on tree saplings are susceptible to being killed by hot flames.

The Prairie Peninsula

In the 1950’s, Robert Gordon mapped the vegetation of Ohio during pre-settlement times, using data from the earliest land surveys. On his famous map below, prairie is indicated in yellow. Notice please that there yellow patches are not scattered uniformly across the state, but are in discrete regional patches.

Gordon's Original Vegetation Map

Gordon’s Original Vegetation Map, with prairie in yellow.

Prairies, having soil high in organic matter and lacking trees needing labor-intensive removal, and well-suited for agriculture. Hence, it is primarily conversion to farmland that has caused the elimination of nearly all of the North American prairies.

Prairie farmer

Nearly all of our prairie has been converted to farmland.

In Ohio and elsewhere. There are three principal types of prairie remnants: pioneer cemeteries, railroad rights-of-way, and lightly grazed areas.

Smith Cemetery in Madison County is a beautiful little (1 acre) State Nature Preserve.

Smith Cemetery is a 1-acre prairie remnant.

 Railroads generally maintain a strip of land alongside their tracks that is untouched by the plow. Here’s one in Union County, called Milford Prairie which is also a State Nature Preserve.

Milford PrairieMilford Prairie was once a railroad right-of-way.

Since it’s woody vegetation, including native trees, that constitute a threat to prairies, light grazing that allows grasses to remain, can be a mechanical alternative to fire. In Crawford County, Daughmer Bur Oak Savanna is a grassland recently, supported a small herd of sheep.

Daughmer Bur Oak Savanna.

Daughmer Bur Oak Savanna.

“Savanna” is the term for a grassland with scattered trees. In the tallgrass prairie region, bur oak (Quercus macrocarpa) is the tree most often seen scattered in prairies.

Q. Hey, what’s the coldest tree? A. Bur-oak (brrr-oak?).

Q. What was J.S. Bach’s favorite tree? A. Bur-oak (baroque?).

O.K. Enough with the jokes already. Bur-oak is a beloved and magnificent tree, well known for being the only tree that grows in prairies. Grasslands with scattered trees are called “savannas.” There is a magnificent savanna in Crawford County, the Daughmer Bur-oak Savannah where this species reigns supreme.  Other species of trees, even native ones such as ashes and maples, are considered weeds in prairies. If ever a windstorm were to sweep through Daughmer Savanah and damage trees, the instructions to tree surgeons dispatched to remedy the damage would be very clear: “If it ain’t bur-oak, don’t fix it!.”

bur-oakBur-oak.

In Ohio we have a few regions where prairie is found, shown on the map below.

Ohio Prairie Regions

Ohio Prairie Regions

COMMON TALLGRASS PRAIRIE GRASSES

The “signature grass” of the tallgrass prairie is big bluestem., Andropogon gerardi.

Big bluestem is the signature tallgrass prairie grass.
Its nickname is “turkey-foot,” because its spikelets are in a few spikes.

Indian grass (Sorghastrum nutans) is another robust prairie grass. Its spikelets contain conspicuously awned florets arranged in loose panicles.

Indian grass is a robust prairie grass.

COMMON TALLGRASS PRAIRIE FORBS 1: ASTERACEAE

The aster family is especially well represented in the tallgrass prairie. The signature prairie forb is one of these; it’s prairie dock (Silphium terebinthinaceum). “Dock” is a general term for a plant with especially large leaves and, prairie dock sure fits that description.

prairie dock

Prairie dock is a large-leaved composite.

Grey-headed coneflower (Ratidiba pinnata) is a composite with gray-brown disk flowers situated on an elongate receptacle. The ray flowers are few in number, long, and somewhat reflexed.

gray-headed coneflower

Grey-headed coneflower has small disk flowers on elongate receptacle.

COMMON PRAIRIE FORBS 3: LEGUMES (FABACEAE)

Perhaps because they have nitrogen-fixing bacterial root nodules that assist growth in nutrient-poor soils, members of the fabulous Fabaceae are prevalent in prairies. One intriguing legume is showy tick-trefoil (Desmodium canadense).Tick-trefoil fruits )MOUSEOVER) are distinctive, as the break apert when mature into single-seeded units with tiny hooked bristles that cause them to stick to animals passing by.

MOUSEOVER the IMAGE to see TICK-TREFOIL FRUITSshowy  tick-trefoilShowy tick-trefoil is a prairie legume.

Round-headed bush-clover (Lespedeza capitata) is a wand-like scruffy-looking legume with small whiteish blossoms arranged in confluent head-like clusters.

bush clover

Round-headed bush-clover

COMMON TALLGRASS PRAIRIE FORBS 4: MINTS (LAMIACEAE)

The Lamiaceae (mint family) includes aromatic herbs with square stems and opposite leaves, and flowers with a sympaetalous corolla that is zygomorphic and usually differentiable into upper and lower lips (bilabiate). Wild begamot, Monarda fistulosa, is a lavender-flowered perennial with broad leaves and the flowers in head-like clusters in the uppermost leaf axils.

wild bergamot

Wild bergamot

False dragonhead (also called “obedient plant” because the large flowers,when pushed to one side, stay in place there instead of bouncing back) is a prairie mint.

false dragonhead

False dragonhead.

COMMON TALLGRASS FORBS 5: BUTTERCUPS (RANUNCULACEAE)

The buttercup family is not especially prevalent in prairies, but the few that do occur there are notably late-blooming compared to other plants in this principally springtime family. The anemones have large white flowers, few in number, atop long pedicels abobe a few pairs of oppositely arranged stem leaves. This is Canada anemone (Anemone canadensis).

Canada anemone

Canada anemone

Tall meadow rue (Thalictrum dasycarpum) is a dioecious plant (i.e., having separate male and female individuals) that is wind-pollinated. Note the abundant small apetalous flowers.

tall meadow-rue

Tall meadow-rue is dioecious.
Photos: male on left; female on right.